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Research
Results (shortly summarized) |
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| [SOILS] [CLIMATE]
[FLORA & VEGETATION] [WWF-INITIATIVE] [Cit.
LITERATURE] [CONCENTRATED SUMMARY] |
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Lithology
& Soils |
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Lithostratigraphic Units, Soil
Physics and Chemistry, Soil Units
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Formations:
Hilly (colline) to middle montane belt: mesozoic ophiolite suites (black and green serpentinites); mainly middle montane
belt: mesozoic - tertiary limestone series; (middle) upper montane - subalpine: paleozoic sandstone series.
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Proved Soil Units
(PDF-File: 40K) according
to FAO-UNESCO (1988):
Trends and intensity of soil-forming processes are - especially under current utilization intensities - basically subject
to relief conditions. In the sense of soil genetical defined chronosequences, highly developed soils were found (independent
of relief position!) on all rock formations.
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Dependent on variant rock structures (of the three formations), the
evolutionary progress and depth of soils is different. Particularly mesozoic deep-seated rocks and palaeozoic
sandstones are deeply weathered. In the area under investigation, generally the depth of soils reflects the economical
utilization of the landscape.
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Clay migration (illimerisation) has been detected in all
highly developed soils
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Vegetation poverty as a result of high concentrations of
plant available heavy metal compounds, often described as typical for soils, derived from serpentinite, has not
been detected in the research area.
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Soil Moisture:
Soils derived from limestone are characterized by the highest field
capacity (FC). It is explained by the lowest content of skeleton, the highest percentage of humus, clay and silt. In
contradiction, the lowest field capacity (FC) has been found in soils derived from serpentinite and sandstones. Soils
derived from serpentinite are deeply weathered, very rich in skeleton with high contents of sand, low percentages of
humus in the topsoil (A-horizon), and the lowest available FC.
Soils with the most favorable requirements for evaporation avoiding
water absorption (e.g. fast and advantageous water intake without evaporation loss) during summer with low precipitation
and high evaporation rates, are deeply weathered soils with high water permeability in the surface soil (e.g.
low content of humus and clay), located at northerly exposed and shady slopes. These pronounced preconditions were achieved
faster especially by soils derived from sandstone under less favorable precipitation conditions, than by soils derived
from limestone.
The exceptional soil moisture conditions from July (August) until
middle of September have been detected to be of exclusive relevance for the distribution of the extrazonal vegetation.
The desiccation period in July decreases with increasing altitude. Generally, shallow and eroded soils dry up quickest.
Under the above mentioned conditions shortages in water balance have been observed especially in the hilly landscapes
(colline belt) and the upper montane to subalpine belt.
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Nutrient Supply:
Under aspects of forestry in soils of all rock formations, the long
and short termed nutrient supply can be described as adequate.
Very different pH Values in soil solutions depend
on relief positions, where soils have been developed. Generally, soils of high slopes derived from sandstone are characterized
by very low pH Values. However, pH Values between 5.5 - 6 in the Ah Horizon can be observed also in very well developed
soils (Luvisols and Phaeozems with clay migration) derived from limestone (mostly dolomite) of small sized areas on
northerly exposed slopes of the middle montane belt with high precipitation rates.
The cation exchange capacity (CEC) depend to the pH
Values and is therefore in soils derived from sandstone mostly 'low' to 'middle'. In contradiction, the CEC is high
in soils developed on different limestones with neutral or low alkaline soil solutions.
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A significant relationship between soil physical and chemical
site conditions and vegetation distribution has been found ONLY in the area of the sandstone formation.
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For the establishment and distribution of plants
of Euxinian origin generally, the soil water availibility during the hot summer season (with low precipitation rates
and extremely high evaporation) has to be interpreted as the most important ecological factor. Thereby highly developed
soils in the sense of soil genetical defined chronosequences are of greater importance than their parent rocks. Nevertheless,
favorable soil conditions have obviously been attained faster by soils derived from sandstone than from serpentinite
and limestone.
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For documentation of Soil Units (cf. TEMap
3) according to FAO-UNESCO a 'Thematical Ecological Map' was developed. |
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Climate
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Overview of the Climatological
Results
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Precipitation:
Precipitation
conditions at the westerly exposed slopes of the Amanus Mtn. are generally determined by the different seasonal macro-climatic
weather. Climatologically the mountain ranges are effected mainly by southerly roaming low pressure areas during
winter season and etesian winds (a drift of air masses between a low pressure area located between the Pontus and Toros
ranges bordering the anatolian highland and a high pressure area above the eastern Mediterranean Sea) during summer
season. The originally dry etesian winds - called Meltemi in Turkey - of the northeast Mediterranean enriches its water
vapor on its way from west to east along the coast and obtains a monsoonal character. The north-south stretching Amanus
Mtn. range is damming up these damp humid air masses with the result of frequent cloudbank formations (cf.
satellite images) (PDF-File: 306K)
during summer time with frequent precipitation events of very different intensities.
The amount and pattern of precipitation is varies extremely from year to year, but especially during summer seasons.
Under the influence of the above mentioned seasonal different precipitation conditions, the easterly (!)
exposed slopes are precedented during winter season. In contradiction, under the influence of the vapor rich etesian
winds during late spring, summer and early autumn, nearly exclusively the westerly exposed slopes enjoy high precipitation
rates.
The coastal plain of the Dörtyol area show already a more or less humid climate. And the transition range from
the hilly landscape to the upper montane belt belongs completely to humid to the per-humid type of bioclimate. Moreover,
the complete montane and subalpine belt is to be added to the per-humid type of bioclimate according to EMBERGER
(1955). Therefore the investigated western slopes of the Amanus Mtn. above Dörtyol show typical humid altitudinal
belts.
Summarizing it can be concluded that the actual Extrazonal Vegetation of the investigated slopes with extremely
strong affinities to Balkan and Pontos vegetation types owes its existence nearly exclusively to precipitation conditions
caused by special effects of typical mediterranean (!) etesian winds of the summer season.
The average precipitation amount of the coastal plain around Dörtyol is quoted with 1.022 mm/y. During the period
of data collection, the following yearly means were measured at different altitudes: 1.300 mm (500 m a.s.l.), 2.300
mm (950 m a.s.l.), 1.800 mm (1.600 m a.s.l.) and 1.300 mm (2.100 m a.s.l.)
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Global Radiation:
An extreme decrease of global radiation during summer season caused by cloudbank
formations (cf. Decrease of Global Radiation
PDF-File: 26K) - especially in the middle montane belt - was observed. In comparison with (theorectically) cloudless
sky, the average reduction of global radiation under conditions of cloudbank formations is more than 40%.
Under consideration of inclination, exposition and mean values of cloud cover, highly significant correlations have
been found between the distribution of vegetation units resp. species populations and the altitudinal gradient of global
radiation.
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Temperature:
(PDF-File: 128K)
The yearly mean altitudinal temperature decrease is 0.61 °C/100 m (200 cm above ground). This value is changing
seasonally and not linear on the altitudinal gradient. During summer season, one can observe a very steep gradient between
the coastal plain and middle montane belt, which can be interpreted as a result of reduced global radiation values effected
by cloudbank formations.
The gradient of temperature between the soil surface and 200 cm above ground is getting steeper with increasing
altitude as a result of an increasing surface heating.
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Flora
& Vegetation |
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Presentation of floristical,
geobotanical and dynamic aspects of vegetation distribution
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Spectrum of Geoelements:
610
species (PDF-File: 238K)
were proved for the research area (incl. 13 species
named by AKMAN [1973] for the slopes above Dörtyol).
NEW:
The Web Project
"Plant Species / Families
of the Amanos Mtn., NW-Slopes above Dörtyol (SE-Turkey, Hatay Province) coll. & det. in the Framework of
the Research Project LöKAT" has been launched in 2011 March, incl. Herbarium
Specimen (digitized) and a great number of Plant Species Photographs.
Geobotanically, the middle montane belt nowadays has
proportionally nearly 44% species of Euro-Siberian origin. In the upper montane belt, this portion rises to more than
63%. The altitudinal distribution of geoelements is determined by the above mentioned unique climate conditions. Cloud
bank formations as regulators of radiation balance during summertime have to be interpreted as the climate variable
with the highest value of explanation. In addition, the relief exposition of species sites is of great importance (cf.
exposition of sample sites
PDF-File: 488K)
However, present abundance and dominance of species
and their populations are determined essentially by anthropo-zoogenous impacts of different intensities (called 'hemerobic
degrees' according to JALAS (1953) and SUKOPP (1968), e.g. woodland
pasture, wood consumption for different purposes and the above mentioned infrastructural development resulting in soil
erosion). In has been observed that habitat changes caused by extensive and intensive economical influence very often
resulted in soil erosion with inauspicious disadvantages of the soil water balance. Especially Euro-Siberian geoelements
were pushed back and their reestablishment disabled. At least euryvalent low-grade east-mediterranean species with wide
amplitudes of growing conditions prevail.
The Mean Total Cover Values
of Geoelements for Different Layers and Altitudes (PDF-File:
54K) were recognized as an important indicator of hemeroby.
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Life Form Spectra:
Independent of plantgeographical origin of species concerned, life
form spectra altitudinally change continuously until the upper timber line. An interference of soils and their variable
moisture supply has not been detected. The most distinctive influence on life form spectra of lower (!) vegetation layers
were caused by (woodland) grazing. As a result of heavy overgrazing, an outstanding quantity of therophytes and hemi-cryptophytes
are growing on limestone soils of the middle montane area with its potential moderate wintertight Euxinian broadleaf-deciduous
mixed forest.
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Growing Periods:
Evaluations
based exclusively on the phenology of Euro-Siberian floral elements have shown the conspicuous temporal parallelism
of growing periods to their far northerly resp. north-westerly located zonal areas. This is especially true for
temporal aspect changes of different species within one altitudinal belt.
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Locations with Similar Climatic Conditions:
(PDF-File: 48K)
Climatologically,
the middle to upper montane belts of the area under investigation show particular similarities with coastal landscapes
of the Black Sea region and central Balkan peninsula, and especially to the middle montane belt with landscapes of the
Dalmatinian and Albanian coast with Quercus cerris and Carpinus orientalis. Merely the per-humid subalpine
belt shows floristical (climatological data are not available) similarities with the subalpine flora and vegetation
of the Pontus Mtn., humid eastern Toros Mtn., mountains of central Greece, and in some respect, of central Italy.
The most convincing conformities concerning the climatic conditions (and composition of flora and vegetation) doubtlessly
exist in the south-eastern area of Black Sea coastal mountains. Even soil types are very similar.
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Syntaxonomical determined Vegetation Units (a Comparison):
A comparison of vegetation units of this project - explicitly not
of synthetic origin -, but as a result of synoptical classification, with traditional syntaxonomical substantiated
'plant communities' of SE-Turkey or E-Mediterranean and Black Sea coastal landscapes has been attempted. Generally,
the comparability lacks reliable data regarding habitat conditions, typical for 'synthetic' phytosociological tables.
The hilly landscapes with strongly degraded vegetation above Dörtyol
show similarities with comparable coastal mountains of Lebanon and Syria, although the climatic conditions are very
different.
Vegetation units of the middle montane belt with Carpinus orientalis
and Quercus cerris show the highest accordances with syntaxonomical orders and alliances with the central Balkan
Peninsula with relatively humid conditions and Pontic coastal ranges with relatively dry conditions and strong human
impact.
Vegetation units of the per-humid middle to upper montane belt,
dominated by Fagus orientalis, show good conformities with beech forests of the Pontus range, Caucasian mountains
and especially with mountains of central Greece. Floristically, the heterogeneous 'beech forest units' are best comparable
with the syntaxonomical determined 'Fagetalia' of SE-Europe.
Of special interest is the floristical composition of the Laurus-Tilia
- unit, restricted to deep shady and moist valleys. A floristical amazingly similar 'plant community' was described
as a pontic-euxinian Buxus-Staphylea pinnata - community, a typical ravine forest of Black Sea coastal beech
forests.
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Geobotanical Relevance of the Amanus Mtn. and Distribution of Endemics:
The extraordinary and extensively extrazonal vegetation of Euxinian origin represents the most homogeneous,
species-rich and southernmost external outpost of typical temperate zonobiomes. Near the Syrian border, this vegetation
reminds strongly of SE-European broadleaf-deciduous forests (bare in winter). Euro-Siberian floral elements clearly
dominate the vegetation of the complete montane belts up to nearly 70%. Already in the colline area of the hilly landscapes,
these geoelements reach 30% of the weighted frequency of all detected species.
As a result of human impact, this vegetation is strongly endangered! It
should be mentioned here that this vegetation is of national and international relevancy and accountable for the most
beautiful mountain landscape of the E-Mediterranean.
This vegetation has been established post-glacial as a consequence of a typical macro climatical summer-weather
constellation of the E-Mediterranean. The species similarity of climatically comparable habitats located in central
or NW-Greece and Black Sea coastal mountains reaches nearly 100%.
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Under consideration of present climate and soil development, an increase of
Euro-Siberian geoelements of Euxinian provenance can without doubt be predicted over a long time span
IF the
present human impact changes to at least oligo-hemerobic habitat conditions. Certainly the most typical Mediterranean
elements would be pushed back from the planar coastal plain and hilly (colline) landscapes. Meso-climatically a zone
with eu-mediterranean vegetation can NOT be expected for the coastal area below the area
under investigation.
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In the research area of LöKAT the percentage of endemic species
reaches only 11.5%. This rate falls short of the value of 30.9%, ascertained by DAVIS (1965-88)
as the average of the 'Flora of Turkey and the East Aegean Islands'. Restricted areas containing endemics are concentrated
on one hand on extreme habitat conditions of the middle montane belt with strong anthropo-zoogenic impact (especially
grazing) and on the other hand on southerly exposed slopes of the subalpine belt with extreme high global radiation
rates.
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'Remote Sensing' and Vegetation Mapping:
The classification of vegetation with Geographical Information Systems (GIS), especially interpretation of satellite
images, in mountainous regions contains numerous incorrect estimates, if images of one season only have been evaluated.
The greatest problems are deviating reflections of vegetation during different growing seasons, resp. phenological conditions
and shading. Within the framework of this research project, two sets of satellite images of different seasons have been
used for the evaluation of vegetation distribution.
In the area under research, three main classes of vegetation were
detected. Regarding the vegetation distribution, images of the growing season have been the most suited. Generally one
can say that multiseasonal images comprising the possibility of the most reliable interpretation of vegetation.
Generally saying, classifications of small-scale mountainous landscapes
are in need of several and serious ground checks and precise knowledge of terrain to be analysed.
Vegetation Units (cf. TEMap
4) were mapped, Abundance und Dominance
of Geoelements represented (cf. TEMap
5), and Vegetation Units compared with the distribution
of Soil Units (3-D-View) (cf. TEMap
6)
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Most
recent initiative regarding the protection and management of the unique Amanos Mtn. - Ecosystem |
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Remarks of Yildiray Lise, WWF - TURKEY on the Amanos Mountains
/ Hatay: ".... However, there are threats over the area: Population is rapidly growing in the region and the
pressure is expanding from towns towards adjacent forests. Amanos Mountains should be legally put under protection as
a National Park, since Natural Park covers wider areas than Nature Reserves and allows zoning for different purposes.
The impact of protection status on indigenous people and the social needs of nearest towns should be carefully evaluated
and an appropriate zoning system has to be established, enabling sustainable use in buffer zones and strict protection
in ecologically important areas. Participatory approach should be applied and a collaborative management should be sought
during the preparation of the management plan."
NOTE:
In the framework of the "Twinning
Project TR02-EN-01 - Capacity Building in the Field of Environment for Turkey, Component 3: Nature",
some additional floristical details on the Amanos Mtn. (Nur
and Daz Daǵları) have been published by
the author of this website.
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Cit.
Literature: |
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AKMAN, Y. (1973a) Contribution à
l'étude de la flore des montagnes de l'Amanus (I-III).- Communications de la Faculté des Sciences de l'Université
d'Ankara, Serie C, Tome 17 C: 1-70. (List of Species)
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AKMAN, Y. (1973b) Aperçu préliminaire sur les conditions
phyto-écologiques de la chaine de l'Amanus dans la region du Hatay (I-III).- Communications de la Faculté des
Sciences de l'Université d'Ankara, Serie C, Tome 17 C: 75-164. (Bioclimate, Soils, Flora)
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AKMAN, Y. (1982) Climats et bioclimats méditerranéens
en Turquie.- In: QUEZEL (ed.) Définition et localisation des écosystémes méditerranéens terrestres,
Colloque de Saint-Maximim (France), 16.-20.11.1981.- Ecologia Mediterranea (Marseille), Tome VIII, Fasc. 1/2: 73-87.
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BERTOVIC, S. (1970) Sumskovegetacijska producja i njihovi
klimatski odnoso kai osnova za regionalnu tipolosku klasifikaciju suma u Hrvatskoj.- Dissertation, Zagreb, Mskr.
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BRAUN-BLANQUET, J. (1964) Pflanzensoziologie (3.Aufl.).- Springer
Verlag, Wien - New York. (865 pp)
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DAVIS, P.D. (1965-1988) (ed.) Flora
of Turkey and the East Aegean Islands.- University Press, Edinburgh. (10 Vol.)
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EMBERGER, L. (1955) Une classification
biogéographique des climats.- Rev. Trav. Lab. Bot. Fac. Sci. 7: 3-43. (Montpellier)
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FAO - UNESCO (1988) Soil Map of the
World, Revised Legend.- World Soil Resources Report 60, Rom.
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HORVAT, I., V. GLAVAC & H. ELLENBERG
(1974) Vegetation Südosteuropas.- Gustav Fischer Verlag, Stuttgart. (768 pp)
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JALAS, J. (1953) Hemerokorit ja hemerobit.-
Luonnon Tutkija 57: 12-16.
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SUKOPP, H. (1968) Der Einfluss des
Menschen auf die Vegetation und zur Terminologie anthropogener Vegetationstypen.- In: TÜXEN (ed.) Pflanzensoziologie
und Landschaftsökologie.- Bericht über das Internationale Symposium in Stolzenau/Weser 1963 der Internationalen
Vereinigung für Vegetationskunde, pp. 65-74.- Verlag Dr. W. Junk N.V., Den Haag.
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VAN DER MAAREL, E. (1979) Transformation of cover-abundance
values in phytosociology and its effects on community similarity.- Vegetatio 39 (2): 97-114.
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© Harald
Kehl all rights reserved TU-Berlin · Fac.VI· Inst. of Ecology Ackerstr. 71-76 D-13355 Berlin
Last updated on
18.10.2011
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